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The rules that allow for networks to exhibit behaviors like those of grid cells have no equivalent in the function of biological brains.
The networks that do not exhibit behaviors like those of grid cells were nonetheless programmed with rules that had proven useful in earlier neural-network studies.
Neural networks can often accomplish tasks that biological brains do, but they are typically programmed with rules to model multiple types of brain cells simultaneously.
Once a neural network is programmed, it is trained on certain tasks to see if it can independently arrive at processes that are similar to those performed by biological brains.