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Macaques in the study called out to and gestured more frequently at humans holding food than at empty-handed humans.
In 2018, researchers who had observed macaques in India's Bandipur National Park calling out to and gesturing at humans designed a study.
The researchers hoped to find out if the macaques were intentionally communicating to try to persuade humans to share their food.
The researchers studied how macaques behaved around both humans holding food and empty-handed humans.